{"id":23434,"date":"2021-01-10T19:37:00","date_gmt":"2021-01-10T23:37:00","guid":{"rendered":"https:\/\/www.shortform.com\/blog\/?p=23434"},"modified":"2021-01-15T17:30:12","modified_gmt":"2021-01-15T21:30:12","slug":"stable-populations","status":"publish","type":"post","link":"https:\/\/www.shortform.com\/blog\/stable-populations\/","title":{"rendered":"Stable Populations and Behavioral Strategies"},"content":{"rendered":"\n<p>How do animals maintain stable populations? Why is ensuring stability important for evolution?<\/p>\n\n\n\n<p>Populations with uncontrolled growth or insignificant growth are not stable populations. Certain behavioral strategies help promote stability through reproduction and choosing when to compete with others.<\/p>\n\n\n\n<p>Read more about stable populations and the behavioral strategies that can help protect stability.<\/p>\n\n\n\n<!--more-->\n\n\n\n<h2 class=\"wp-block-heading\">Ensuring Stable Populations<\/h2>\n\n\n\n<p><strong>How animals decide when to reproduce is a complicated question. <\/strong>Some scientists, most notably zoologist Vero Copner <a href=\"https:\/\/www.shortform.com\/blog\/theories-of-reproduction\/\">Wynne-Edwards<\/a>, believe that animals control their reproduction rates to avoid overpopulating an area. They&#8217;re aiming for stable populations. However, this would be a group-selection based theory, which doesn\u2019t seem to mesh with the idea of the selfish gene. It\u2019s worth noting that Wynne-Edwards was a prominent champion of the group-selection theory of evolution, and his ideas are influential enough to be worth considering\u2014even if only to refute them.<\/p>\n\n\n\n<p>Whatever the reason, it\u2019s clear that animal populations don\u2019t grow at the incredible rates they\u2019d be theoretically capable of. Many populations remain fairly stable, with birth and death rates roughly matching each other. Others, like lemmings, grow rapidly and then decline sharply, sometimes to the point of total extinction in a particular area.<\/p>\n\n\n\n<p>Starvation is one major factor in keeping animal populations under control. If animals reproduce too quickly for an area to support, naturally many of them will starve. <strong>However, starvation cannot fully explain how animal populations stay under control. <\/strong>If starvation were the only control set on population size, scientists would expect all creatures to work like lemmings do: Their numbers would increase exponentially until the region couldn\u2019t support them, then suddenly crash as most of them starved.&nbsp;<\/p>\n\n\n\n<p>Therefore, it\u2019s clear that there are methods limiting birth rates as well as death rates\u2014animals don\u2019t have infinite numbers of offspring. <strong>The question is not <\/strong><strong><em>whether <\/em><\/strong><strong>birth rates are controlled, but <\/strong><strong><em>why <\/em><\/strong><strong>they are controlled.<\/strong><\/p>\n\n\n\n<p>The group-selection theory would say the reason is altruistic: Animals regulate their birth rates for the good of the entire population. The <a href=\"https:\/\/www.shortform.com\/blog\/selfish-gene-theory\/\">selfish gene theory<\/a> would say that it\u2019s selfish: Animals regulate their birth rates because it gives them and their offspring the best <a href=\"https:\/\/www.shortform.com\/blog\/premature-birth-chances-of-survival-at-weeks-24-26\/\">chance of survival<\/a>.<\/p>\n\n\n\n<h3 class=\"wp-block-heading\"><strong>Game Theory and Behavioral Strategies<\/strong><\/h3>\n\n\n\n<p>Game theory can be used to explain behaviors, especially at the population level. By using game theory, scientists can determine effective behavioral strategies. They can also find\u2014or at least approximate\u2014an ESS for a population made up of such behaviors. These behavioral strategies support stable populations.<\/p>\n\n\n\n<p><strong>The Prisoner\u2019s Dilemma is a logistical riddle closely tied to game theory. <\/strong>In the Prisoner\u2019s Dilemma there are two players, each with two options: Cooperate and Betray. Neither player knows which option the other has chosen, and they are not allowed to influence the other\u2019s choice in any way.<\/p>\n\n\n\n<p>If both players choose Cooperate, they each get a significant payout\u2014but a smaller one than in the next situation. If one player Cooperates and the other Betrays, the betrayer gets a large payout while the cooperative player suffers a large penalty. If both players Betray, they each suffer a small penalty. Cooperation and betrayal are altruistic and selfish actions, respectively. Therefore, everything we know about the Dilemma could be compared to nature.<\/p>\n\n\n\n<p><strong>In any single instance of the game, the logical choice is to simply pick Betray. <\/strong>If your opponent chose Cooperate, you\u2019ll get a larger payout than if you\u2019d also chosen Cooperate. If your opponent chose Betray, you\u2019ll suffer a smaller penalty than if you\u2019d chosen Cooperate.<\/p>\n\n\n\n<p><strong>However, the strategy becomes much more complex if you play over and over again.<\/strong> Now one Always-Betray strategy can get stuck in a penalizing loop with another one, while more cooperative strategies have the chance to reap mutual benefits. Of course, an Always-Cooperate strategy will still lose every time to an Always-Betray strategy.&nbsp;<\/p>\n\n\n\n<h3 class=\"wp-block-heading\"><strong>Which Strategies Are the Most Effective?<\/strong><\/h3>\n\n\n\n<p>Professor <a href=\"https:\/\/www.shortform.com\/blog\/robert-axelrod\/\">Robert Axelrod<\/a> once invited programmers to create programs that use Prisoner\u2019s Dilemma strategies, which he then entered into a virtual \u201ctournament.\u201d In this tournament, each program played 200 rounds of Prisoner\u2019s Dilemma with each of the others, including a copy of itself. Whichever program had the highest total score at the end was the winner.<\/p>\n\n\n\n<p>There were a number of complex strategies submitted for this tournament, some of which were quite cutthroat. However, the surprising winner of the tournament was a simple program called Tit for Tat. This program always played Cooperate on the first round, then for each round after that it simply copied what its opponent had done last. This made it so cooperative strategies were rewarded, while aggressive strategies were punished. Afterward, Axelrod calculated that a so-called \u201cTit for Two Tats\u201d program would have done even better; such a program wouldn\u2019t Betray until having been betrayed twice in a row itself, and would therefore have avoided some penalizing loops that Tit for Tat got caught in.&nbsp;<\/p>\n\n\n\n<p><strong>However, Tit for Tat alone\u2014or any program that never Betrays first\u2014can\u2019t be considered a true ESS.<\/strong> Such a population could be infiltrated by a mutant program that <em>never <\/em>picks Betray. As long as there are no aggressive programs in the field, that new program can spread. That would leave the entire population vulnerable if an aggressive program were later introduced. Axelrod coined the term \u201cCollectively Stable Strategy\u201d to describe this situation (remember that an ESS <em>cannot <\/em>be invaded by another strategy).<\/p>\n\n\n\n<h3 class=\"wp-block-heading\"><strong>Attaining Stab<\/strong>le Populations<\/h3>\n\n\n\n<p><strong>While it will be difficult for another strategy to invade a population of Always-Betray individuals, it\u2019s almost inevitable that it will happen eventually. <\/strong>A population of creatures with the Always-Betray strategy will continually weaken itself, while a population of more cooperative individuals will prosper and spread. Sooner or later the balance will tip back toward the cooperative population.<\/p>\n\n\n\n<p>Therefore, while Always-Betray is technically an <a href=\"https:\/\/www.shortform.com\/blog\/evolutionarily-stable-strategy\/\">Evolutionarily Stable Strategy<\/a> (in the sense that no other program could do better in a population of Always-Betray), and Tit for Tat is technically not an ESS (because it could be invaded by a mutant program and eventually overthrown), it could be said that Tit for Tat has a long-term stability that Always-Betray lacks.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>How do animals maintain stable populations? Why is ensuring stability important for evolution? Populations with uncontrolled growth or insignificant growth are not stable populations. Certain behavioral strategies help promote stability through reproduction and choosing when to compete with others. Read more about stable populations and the behavioral strategies that can help protect stability.<\/p>\n","protected":false},"author":6,"featured_media":390,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_jetpack_memberships_contains_paid_content":false,"footnotes":""},"categories":[42,21,24],"tags":[187],"class_list":["post-23434","post","type-post","status-publish","format-standard","has-post-thumbnail","hentry","category-motivation","category-philosophy","category-society","tag-the-selfish-gene","","tg-column-two"],"yoast_head":"<!-- This site is optimized with the Yoast SEO Premium plugin v24.3 (Yoast SEO v24.3) - https:\/\/yoast.com\/wordpress\/plugins\/seo\/ -->\n<title>Stable Populations and Behavioral Strategies - Shortform Books<\/title>\n<meta name=\"description\" content=\"To ensure evolutionary success, animals want stable populations. See how they pursue reproduction and other strategies for stability.\" \/>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/www.shortform.com\/blog\/stable-populations\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Stable Populations and Behavioral Strategies\" \/>\n<meta property=\"og:description\" content=\"To ensure evolutionary success, animals want stable populations. See how they pursue reproduction and other strategies for stability.\" \/>\n<meta property=\"og:url\" content=\"https:\/\/www.shortform.com\/blog\/stable-populations\/\" \/>\n<meta property=\"og:site_name\" content=\"Shortform Books\" \/>\n<meta property=\"article:published_time\" content=\"2021-01-10T23:37:00+00:00\" \/>\n<meta property=\"article:modified_time\" content=\"2021-01-15T21:30:12+00:00\" \/>\n<meta property=\"og:image\" content=\"https:\/\/www.shortform.com\/blog\/wp-content\/uploads\/2019\/07\/factfulness-crowd.jpg\" \/>\n\t<meta property=\"og:image:width\" content=\"823\" \/>\n\t<meta property=\"og:image:height\" content=\"433\" \/>\n\t<meta property=\"og:image:type\" content=\"image\/jpeg\" \/>\n<meta name=\"author\" content=\"Rina Shah\" \/>\n<meta name=\"twitter:card\" content=\"summary_large_image\" \/>\n<meta name=\"twitter:label1\" content=\"Written by\" \/>\n\t<meta name=\"twitter:data1\" content=\"Rina Shah\" \/>\n\t<meta name=\"twitter:label2\" content=\"Est. reading time\" \/>\n\t<meta name=\"twitter:data2\" content=\"5 minutes\" \/>\n<script type=\"application\/ld+json\" class=\"yoast-schema-graph\">{\"@context\":\"https:\/\/schema.org\",\"@graph\":[{\"@type\":\"Article\",\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/#article\",\"isPartOf\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/\"},\"author\":{\"name\":\"Rina Shah\",\"@id\":\"https:\/\/www.shortform.com\/blog\/#\/schema\/person\/d07435826f13a8f422d5d333dceac287\"},\"headline\":\"Stable Populations and Behavioral Strategies\",\"datePublished\":\"2021-01-10T23:37:00+00:00\",\"dateModified\":\"2021-01-15T21:30:12+00:00\",\"mainEntityOfPage\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/\"},\"wordCount\":1037,\"commentCount\":0,\"publisher\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/#organization\"},\"image\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/#primaryimage\"},\"thumbnailUrl\":\"https:\/\/www.shortform.com\/blog\/wp-content\/uploads\/2019\/07\/factfulness-crowd.jpg\",\"keywords\":[\"The Selfish Gene\"],\"articleSection\":[\"Motivation\",\"Philosophy\",\"Society\"],\"inLanguage\":\"en-US\",\"potentialAction\":[{\"@type\":\"CommentAction\",\"name\":\"Comment\",\"target\":[\"https:\/\/www.shortform.com\/blog\/stable-populations\/#respond\"]}]},{\"@type\":\"WebPage\",\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/\",\"url\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/\",\"name\":\"Stable Populations and Behavioral Strategies - Shortform Books\",\"isPartOf\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/#website\"},\"primaryImageOfPage\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/#primaryimage\"},\"image\":{\"@id\":\"https:\/\/www.shortform.com\/blog\/stable-populations\/#primaryimage\"},\"thumbnailUrl\":\"https:\/\/www.shortform.com\/blog\/wp-content\/uploads\/2019\/07\/factfulness-crowd.jpg\",\"datePublished\":\"2021-01-10T23:37:00+00:00\",\"dateModified\":\"2021-01-15T21:30:12+00:00\",\"description\":\"To ensure evolutionary success, animals want stable populations. 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